I presume as you have received an email mentioning the fitment of parts requiring a headspace check, then you are involved in your rifle in more depth and detail than you usually are, you definitely need to get a qualified gunsmith to fit said parts which includes a headspace check.
Now to answer your question in a monotone singular syllable prose
When ammo is chambered in your rifle (ready to fire) it is sitting in there and held quite snug, (but not as tight as Dundee clinging to a stubbie) there are a few thousandths of an inch clearance to ensure the ammo can enter and exit the rifles chamber either fired or unfired.
Too little clearance and there are a few problems which appear quite quickly, too much clearance and again problems appear, some quite serious.
There are other important reasons the ammo for your rifle needs this clearance but we have a good head start here.
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